The metabolic cost of walking on gradients with a waddling gait

SUMMARY Using open-flow respirometry and video footage (25 frames s–1), the energy expenditure and hindlimb kinematics of barnacle geese, Branta leucopsis, were measured whilst they were exercising on a treadmill at gradients of +7 and –7 deg, and on a level surface. In agreement with previous studies, ascending a gradient incurred metabolic costs higher than those experienced on level ground at comparable speeds. The geese, however, are the...